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June 7, 2016; 113 (23)

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Cover image: Pictured are a maxilla and mandible of Australopithecus deyiremeda, along with a hypothesized cranial outline, and the 3.3 million-year-old Burtele partial foot from the Woranso-Mille, central Afar, Ethiopia. The Burtele foot has an opposable big toe, unlike the human-like foot inferred for the contemporary hominin Australopithecus afarensis, Lucy's species. In a Perspective in the Human Origins Special Feature, Yohannes Haile-Selassie et al. discuss how fossils such as these provide compelling evidence for the coexistence of more than one hominin species during the middle Pliocene, 3.3–3.5 million years ago. However, the small numbers of specimens from each species complicate our understanding of the relationships between these early hominin species. See the Perspective by Haile-Selassie et al. on pages 6364–6371. Image courtesy of Yohannes Haile-Selassie.

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